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MiddleLayer

Jobs at startups backed by the best investors.

We only list companies backed by top investors, and roles taken from their real career pages.

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© 2026 MiddleLayerIndependent · not affiliated with YC or a16z

MiddleLayer

Jobs at startups backed by the best investors.

We only list companies backed by top investors, and roles taken from their real career pages.

Jobs Firms Companies How it works API

© 2026 MiddleLayerIndependent · not affiliated with YC or a16z

8,522roles

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Startup Partnerships LeadPosted Jun 5

AnthropicSan FranciscoSalesPrivateSequoiaSpark

Staff + Sr. Software Engineer, Cloud Inference Launch EngineeringPosted Jun 3

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff + Sr. Software Engineer, Cloud InferencePosted Jun 3

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Security EngineerPosted Feb 13

AnthropicSan FranciscoSecurityPrivateSequoiaSpark

Staff+ Software Engineer, Enterprise Knowledge WorkPosted Jul 2

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Safeguards Review ToolingPosted Jul 13

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Safeguards ML InfrastructurePosted Jun 24

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Safeguards Evals Posted Jun 9

AnthropicSan FranciscoSafeguards (Trust & Safety)PrivateSequoiaSpark

Staff+ Software Engineer, SafeguardsPosted Oct 14

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Public SectorPosted May 4

AnthropicRemoteEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, PlatformPosted Apr 8

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff Software Engineer, People ProductsPosted Feb 13

AnthropicRemoteEngineeringPrivateSequoiaSpark

Staff Software Engineer, Labs: Applied AIPosted Jul 8

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff Software Engineer, iOSPosted Aug 28

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Inference RuntimePosted Jun 12

AnthropicRemoteEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Identity & Access ControlsPosted Jun 18

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, GRC PlatformPosted Jun 8

AnthropicSan FranciscoSecurityPrivateSequoiaSpark

Staff+ Software Engineer, Full-stackPosted Apr 8

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Financial FraudPosted Jul 10

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, ExperimentationPosted Jul 2

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, EnterprisePosted Jul 1

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Developer ProductivityPosted Feb 13

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Developer AccelerationPosted Jul 2

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Data InfrastructurePosted Feb 13

AnthropicRemoteEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, DatabasesPosted Jul 7

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Claude App InfrastructurePosted Jan 13

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff Software Engineer, Front-EndPosted Dec 18

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Capacity EngineeringPosted Jul 14

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, CachingPosted Jul 7

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff Software Engineer, Billing PlatformPosted Mar 10

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, BackendPosted Apr 8

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff Software Engineer, AI ReliabilityPosted Feb 7

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Software Engineer, Account AbusePosted Feb 24

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff + Senior Software Engineer, Inference DeploymentPosted Jun 29

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff + Senior Software Engineer, InferencePosted Jun 8

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Security Engineer, Risk EngineeringPosted Jun 8

AnthropicSan FranciscoSecurityPrivateSequoiaSpark

Staff Research Engineer, Discovery TeamPosted Apr 23

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff Engineer, Datacenter Server LifecyclePosted Mar 11

AnthropicSan FranciscoEngineeringPrivateSequoiaSpark

Staff+ Application Security Engineer - M&A Posted Jul 10

AnthropicRemoteSecurityPrivateSequoiaSpark

Staff+ Application Security EngineerPosted Jan 22

AnthropicRemoteSecurityPrivateSequoiaSpark

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8,522 open roles